1. Прикладна мікробіологія та біотехнологія (метаболічна інженерія, аналітична біотехнологія).
2. Біохімія (регуляторні аспекти метаболізму в мікроорганізмах).
3. Білкова біотехнологія (виділення та очищення білків, їх структурна та фізико-хімічна характеристика).
4. Ензимологія (фундаментальні та прикладні аспекти), Біосенсорика, Екологічна біотехнологія, Нанобіотехнологія.

Автор понад 560 наукових публікацій, у тому числі 215 повноформатних робіт та 12 винаходів (більше 125 – у журналах з імпакт-фактором). Індекс Хірша (h-index) публікацій ⊄ 24 (Scopus).

Список публікацій

2000-present – Head of the Department of Analytical Biotechnology, Institute of Cell Biology, NAS of Ukraine;

1985-2000 – Senior Researcher, Institute of Cell Biology, NAS of Ukraine, Lviv, Ukraine;

1980-1985 – Assistant of Professor (Docent), Chair of Biochemistry, Ivan Franko State University, Lviv, Ukraine;

1978-1980 – Senior Researcher, Institute of Hematology and Blood Transfusiology, Lviv, Ukraine.

1974-1977 – Postgraduate student of Moscow University, Chemical Faculty, Department of Chemistry of Natural Compounds;

1969-1974 – Student of the Moscow State University, Chemical Faculty, Department of Chemistry of Natural Compounds.

2005 – Professor, Diploma on Genetics and Biotechnology, Ivan Franko National University of Lviv;

2001 – DrSc. (habilitation) Thesis "Pathways of energy supply and detoxification in methylotrophic yeasts and their directed modification for development of enzymatic and biosensor analytical systems". Speciality: Biochemistry;

1983 – Associated Professor (Docent) Diploma on Biochemistry, Ivan Franko State University of Lviv;

1978 – Ph.D Thesis "Multiple molecular forms of equine pepsin", Moscow State University. Speciality: Bioorganic Chemistry, Chemistry of Natural and Biologically Active Compounds.

2012 (October-November) – Wroclaw University (Poland), visiting professor;

2000 (August–September) – Leeds and Oxford Universities (United Kingdom);

1996 (November-December) – Florence University (Italy), Chemical Department;

1995 (September-October) – Florence University (Italy), Chemical Department;

1991 (October-November) – Wroclaw University (Poland), Institute of Microbiology;

1985 (February-June) – Moscow State University, Biological Faculty, Department of Biochemistry.

2018 – Winner of the State Prize of Ukraine in Science and Technology;

1993 and 1998 – Awards of Soros International Science Foundation.

2008–2016 – University of Rzeszow (Poland), Department of Biotechnology, Professor;

2006–2007 – Ivan Franko Pedagogical University of Drohobych, Professor;

2002–2005 – Ibid, Department of Genetics and Biotechnology, Professor;

1987–1997 – Ibid, Department of Genetics and Biotechnology, Associated Professor (Docent);

1980–1985 – Ivan Franko State University of Lviv, Department of Biochemistry, Associated Professor (Docent).